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Polarized M1 macrophages are effector cells induced by pro-inflammatory cytokines, such as tumour necrosis factor α (TNF-α) and interferon-γ (IFN-γ) or by other pro-inflammatory stimuli including bacterial lipopolysaccharide (LPS), [60,61]

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Factors that cause sperm DNA fragmentation (SDF) are complex, and among these factors, old age or exposure to toxic substances can be mentioned [105], [106]