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Investigation of tissue specific expression of GTRs with the thorough analysis of loss-of-function mutants suggested the role of GTR2 in GSL phloem loading, whereas GTR1 is assisting GTR2 in phloem loading but expected to be additionally involved in distributing GSL within the leaf ( in vivo , GTR1 and GTR2 are likely to also be involved in bidirectional distribution of GSLs between the roots and rosettes, substantiating the role of both phloem and xylem as their transport pathways ( The NRT/PTRs constitute a large plant transporter family belonging to the ubiquitous POT (Tsay et al., 2007)

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