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For P134L, it is probable that the substitution of this proline (a turn-inducing residue) with leucine (a non-turn-inducing residue) affected the conformation of the protein backbone, thus impairing (but not preventing) hydrogen bonding between the NH 2 of nearby Arg 132 and the oxygen atom of the 3’-phosphate in PAPS 20,28

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Lancet 2, 1300–1304 (1987)